Andrea H. Lloyd
Assistant Professor
Department of Biology
Middlebury College
Middlebury, VT 05443
LLOYD@MIDDLEBURY.EDU
fax:802-443-2072
phone:802-443-3165
Fall 2003 Office Hours (Bicentennial Hall 372)
Monday 9:00-10:00 am
Friday 10:00 am -12:00 pm
or by appointment
Abbreviated Curriculum vitae
Current Research Projects
Dynamics of black spruce (Picea mariana) at
its northern limit in the Brooks Range, Alaska

This research project, funded by the Bonanza
Creek LTER Program, investigates the dynamics of black spruce forests at
the species' northern limit in the Brooks Range, Alaska. Black spruce forests
are the most widespread forest type in interior Alaska, but black spruce becomes
increasingly less common as treeline is approached, and treeline forests are
dominated almost completely by white spruce.
We are seeking to understand:
- the relative importance of climatic/non-climatic controls
over black spruce distribution
- the fire regime of northernmost black spruce forests
- competitive interactions between white and black spruce at
the northern limit of black spruce.

Black spruce are co-dominant with white spruce near their northern
limit. This site, near milepost 200 on the Dalton Highway, is approximately
60% black spruce and 40% white spruce. Evenly mixed stands like this are much
more unusual in the central part of black spruce's range.
Initial results from this research are summarized in the following
:
Wilson, A. E. 2003. The population dynamics of Picea mariana
at the northernmost range limit in the Brooks Range, Alaska: A matrix population
model. Honors Thesis, Middlebury College, VT. To download a PDF version of this
thesis, click here. (File is approximately 1.2
MB.)
Lloyd, A.H., A. E. Wilson, and C. L. Fastie. 2003. Population
dynamics of black spruce at its northern limit in Alaska. Poster presentation
at the Ecological Society of America Annual Meeting. For link to Abstract, click
here.
Response of the Alaskan boreal
forest to climate change
(Funded by the Bonanza
Creek Long-Term Ecological Research Program and the
National Science Foundation's Land-Atmosphere-Ice
Interactions program in the Office of Polar Programs).
This research project is focused on the following questions:
- Is global warming likely to cause the northward expansion
of boreal forests?
- How do climate and disturbance (especially by fire) interact
to control spruce recruitment at the forest-tundra boundary?
- Has warming in the last 100 years caused changes in the structure/distribution
of treeline forests?

Treeline at Twelvemile Summit, along the Steese Highway.
Visit our outreach
web site, constructed by John Mauro as part of his
senior honors thesis work in 1999.
Recent publications from this project:
- Lloyd, A.H. and C.L. Fastie. 2002. Spatial and temporal variability in tree
growth and climate response of treeline trees in Alaska. Climatic Change.
52:481-509.
- Lloyd, A.H., T. S. Rupp, C.L. Fastie, and A. M. Starfield. (2003). Patterns
and dynamics of treeline advance on the Seward Peninsula, Alaska. Journal
of Geophysical Research-Atmospheres. 108 (D2): 8161, doi: 10.1029/2001JD000852.
- Lloyd, A.H. and C.L. Fastie (2003). Recent changes in treeline forest distribution
and structure in interior Alaska. Ecoscience. 10(2):176-185.
- Lloyd, A.H., K. Yoshikawa, C. L. Fastie, L. Hinzman, and M. Fraver. (2003).
Effects of permafrost degradation on woody vegetation at arctic treeline on
the Seward Peninsula, Alaska. Permafrost and Periglacial Processes. vol. 14.
doi: 10.1002/ppp.446.
Any opinions, findings, conclusions,
or recommendations expressed in the material on this site are those of the author(s)
and do not necessarily reflect the views of the supporting agencies.
Teaching
BI 190, Fall 2002
BI 323, Spring
2003
BI 490,
Spring 2003
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